Police officers were called to the scene just outside the stadium at about 19:20 BST He died in hospital shortly before 21:00 BST.
Six people, all thought to be Wales supporters, were arrested and police do not believe any England fans were involved. The Met's Homicide and Serious Crime Command is investigating.
'Extensive efforts' Emergency services were called to an area outside the sporting venue at about 19:20 BST, following reports of an assault shortly before kick-off.
A London Ambulance Service (LAS) spokesman said: "I can confirm that LAS got a call at 7.20pm to reports of an assault outside Wembley Stadium.
"Staff treated a 44-year-old man who had suffered a cardiac arrest. Extensive efforts were made to resuscitate the patient and he was taken to hospital."
Staff on standby at the international match were on the scene within three minutes, the spokesman said.
The victim was taken to a north London hospital in an ambulance belonging to a private medical contractor employed by Wembley.
A post-mortem examination will be arranged in due course.
Police believe they have identified the victim but await formal confirmation that all his next of kin have been informed.
'No shouting' Ollie Ayton, who was at the match and close by when the incident happened, said he had heard the man fall to the ground.
"I did not see him fall as I was facing the other way but I turned immediately to see him unconscious.
"There did not appear to be any fight before; I cannot confirm this as I did not see it, but there was no excess noise or shouting that signalled a fight.
"My friends and I went to watch the match after we realised we couldn't help.
"I left 85 minutes into the game and there were lots of police officers and the area was cordoned off."
England won the European Championship qualifying match 1-0.
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